Rusty thinks Cappie might be ruining his new relationship with Lana, so he convinces Dale to help him save Cappie from himself. Casey and Ashleigh try to do a mixer with ZBZ and Omega Chi to get the sorority its reputation back.
Cappie and Evan's friendship is investigated by Casey. Rusty goes with Dale and Calvin to a comic book convention as a distraction from Jordan. Ashleigh and Fisher decide to begin dating again.
It's Founders Weekend at the ZBZ House and Casey uses the opportunity to scrap for donations from the alums while Rusty tries to keep Jordan from dropping out of school.
Shocked to learn that ZBZ has slipped in the ranks down to fourth, Casey is determined to bring ZBZ back to the number one spot by motivating their pledge class to take action, believing that, if they create a cool photo shoot for the Lamda Sig calendar, it could improve their house’s ranking. However new pldge Abby convinces the other pledges to boycott the shoot so, in order to make nice with the ZBZ pledges, Ashley and Casey get them all makeovers.
It's Dry Weekend at Cyprus Rhodes University, and Casey plans a non-alcohol party night amongst all of the houses but, when she and Evan are partnered together to patrol each of the houses for signs of any prohibited beverages, they each have a brownie at the Psi Phi house unaware of its secret ingredient, causing them to hide out in the ZBZ bathroom until the effects wear off. Meanwhile, Ashleigh frantically tries to find Casey, and Fisher offers to help, making her further question whether or not she should take him back. Also Grant's high school girlfriend, Whitney surprises Grant andd Calvin by visiting for his birthday.
As Rusty and Jordan are awkwardly deciding when to take their relationship to the next level, the Gotcha Game comes between them as the final two players left in the game. Leading Cappie to become Rusty's bodyguard and Rebecca as Jordan's, as the happy couple battle it out so their house can win the game who will pull the trigger first. Meanwhile, Casey takes over as the ZBZ Panhellenic delegate, butting heads with the president, Katherine, over the annual "Undie Run". After Katherine has the event canceled, Casey uses Evan's help to get enough signatures to overturn the decision and save the traditional event by using it as a fundraising event for charity. And Calvin and Grant are trying desperately to find some alone time so they can be together away from their raucous Omega Chi brothers. In the end, they decide to leave campus for a romantic dinner, but are surprised by who turns out to be their waiter.
It's father-daughter weekend at ZBZ, as tension builds at the house after the secret of whom Rebecca's kissed is revealed. Meanwhile, Rusty tries to impress Jordan's father with sports, and Dale and Cappie explore different religions.
The students at CRU feel the ramifications of the Kappa Tau's "End of the World" Party. Rusty risks being dropped from the honors program if he cannot raise his grade in organic chemistry while Dale faces a huge moral dilemma. Also the ZBZ's celebrates father-daughter weekend, however they've found themselves sliding in the ranks among the other sorority houses, leading Casey, Rebecca, and Ashleigh do everything they can to maintain their status as one of the top houses at Cyprus Rhodes. Meanwhile, after learning of Max’s departure, Cappie wonders if he made the right decision by telling Casey he wasn’t interested in giving their relationship another chance, while Casey struggles with the aftermath of Cappie’s unexpected rejection.
After she falls into a manhole risking her life, coupled with the Kappa Tau's throwing an "End of the World" party, Casey is sent into an emotional tailspin, questioning her relationship with Max and her lingering feelings for Cappie. Meanwhile, Ashleigh faces her own punishment with Pan Hellenic after a complaint is lodged by Frannie for the Homecoming Float sabotage.
As the Homecoming Week float competition heats up, Rusty and Ashleigh work hard to obtain a win while Casey and Cappie try to avoid one another. Elsewhere, Evan learns that he may lose his trust fund, and Ashleigh uncovers a secret of Fisher's.